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New Years Solitaire. Valentine Solitaire. St Patricks Solitaire. Some spiders care for their young, for example a wolf spider 's brood clings to rough bristles on the mother's back,  and females of some species respond to the "begging" behaviour of their young by giving them their prey, provided it is no longer struggling, or even regurgitate food.
Like other arthropods , spiders have to molt to grow as their cuticle "skin" cannot stretch. Spiders occur in a large range of sizes.
The smallest, Patu digua from Colombia, are less than 0. Only three classes of pigment ommochromes , bilins and guanine have been identified in spiders, although other pigments have been detected but not yet characterized.
Melanins , carotenoids and pterins , very common in other animals, are apparently absent. In some species, the exocuticle of the legs and prosoma is modified by a tanning process, resulting in a brown coloration.
Guanine is responsible for the white markings of the European garden spider Araneus diadematus. It is in many species accumulated in specialized cells called guanocytes.
In genera such as Tetragnatha , Leucauge , Argyrodes or Theridiosoma , guanine creates their silvery appearance. While guanine is originally an end-product of protein metabolism, its excretion can be blocked in spiders, leading to an increase in its storage.
The white prosoma of Argiope results from bristles reflecting the light, Lycosa and Josa both have areas of modified cuticle that act as light reflectors.
While in many spiders color is fixed throughout their lifespan, in some groups, color may be variable in response to environmental and internal conditions.
For example, the abdomen of Theridion grallator will become orange if the spider ingests certain species of Diptera and adult Lepidoptera , but if it consumes Homoptera or larval Lepidoptera, then the abdomen becomes green.
Morphological changes require pigment synthesis and degradation. In contrast to this, physiological changes occur by changing the position of pigment-containing cells.
Misumena vatia for instance can change its body color to match the substrate it lives on which makes it more difficult to be detected by prey. Juveniles of some spiders in the families Anyphaenidae , Corinnidae , Clubionidae , Thomisidae and Salticidae feed on plant nectar.
Laboratory studies show that they do so deliberately and over extended periods, and periodically clean themselves while feeding.
These spiders also prefer sugar solutions to plain water, which indicates that they are seeking nutrients.
Since many spiders are nocturnal, the extent of nectar consumption by spiders may have been underestimated. Nectar contains amino acids , lipids , vitamins and minerals in addition to sugars, and studies have shown that other spider species live longer when nectar is available.
Feeding on nectar avoids the risks of struggles with prey, and the costs of producing venom and digestive enzymes. Various species are known to feed on dead arthropods scavenging , web silk, and their own shed exoskeletons.
Pollen caught in webs may also be eaten, and studies have shown that young spiders have a better chance of survival if they have the opportunity to eat pollen.
In captivity, several spider species are also known to feed on bananas , marmalade , milk , egg yolk and sausages. The best-known method of prey capture is by means of sticky webs.
Varying placement of webs allows different species of spider to trap different insects in the same area, for example flat horizontal webs trap insects that fly up from vegetation underneath while flat vertical webs trap insects in horizontal flight.
Web-building spiders have poor vision, but are extremely sensitive to vibrations. Females of the water spider Argyroneta aquatica build underwater "diving bell" webs that they fill with air and use for digesting prey, molting, mating and raising offspring.
They live almost entirely within the bells, darting out to catch prey animals that touch the bell or the threads that anchor it.
Net-casting spiders weave only small webs, but then manipulate them to trap prey. Those of the genus Hyptiotes and the family Theridiosomatidae stretch their webs and then release them when prey strike them, but do not actively move their webs.
Those of the family Deinopidae weave even smaller webs, hold them outstretched between their first two pairs of legs, and lunge and push the webs as much as twice their own body length to trap prey, and this move may increase the webs' area by a factor of up to ten.
Experiments have shown that Deinopis spinosus has two different techniques for trapping prey: backwards strikes to catch flying insects, whose vibrations it detects; and forward strikes to catch ground-walking prey that it sees.
These two techniques have also been observed in other deinopids. Walking insects form most of the prey of most deinopids, but one population of Deinopis subrufa appears to live mainly on tipulid flies that they catch with the backwards strike.
Mature female bolas spiders of the genus Mastophora build "webs" that consist of only a single "trapeze line", which they patrol.
They also construct a bolas made of a single thread, tipped with a large ball of very wet sticky silk. They emit chemicals that resemble the pheromones of moths , and then swing the bolas at the moths.
The spiders eat the bolas if they have not made a kill in about 30 minutes, rest for a while, and then make new bolas. Instead they release different pheromones that attract moth flies , and catch them with their front pairs of legs.
The primitive Liphistiidae , the "trapdoor spiders" of the family Ctenizidae and many tarantulas are ambush predators that lurk in burrows, often closed by trapdoors and often surrounded by networks of silk threads that alert these spiders to the presence of prey.
Some jumping spiders of the genus Portia hunt other spiders in ways that seem intelligent,  outflanking their victims or luring them from their webs.
Laboratory studies show that Portia ' s instinctive tactics are only starting points for a trial-and-error approach from which these spiders learn very quickly how to overcome new prey species.
Ant-mimicking spiders face several challenges: they generally develop slimmer abdomens and false "waists" in the cephalothorax to mimic the three distinct regions tagmata of an ant's body; they wave the first pair of legs in front of their heads to mimic antennae , which spiders lack, and to conceal the fact that they have eight legs rather than six; they develop large color patches round one pair of eyes to disguise the fact that they generally have eight simple eyes, while ants have two compound eyes; they cover their bodies with reflective bristles to resemble the shiny bodies of ants.
In some spider species, males and females mimic different ant species, as female spiders are usually much larger than males. Ant-mimicking spiders also modify their behavior to resemble that of the target species of ant; for example, many adopt a zig-zag pattern of movement, ant-mimicking jumping spiders avoid jumping, and spiders of the genus Synemosyna walk on the outer edges of leaves in the same way as Pseudomyrmex.
Ant mimicry in many spiders and other arthropods may be for protection from predators that hunt by sight, including birds, lizards and spiders.
However, several ant-mimicking spiders prey either on ants or on the ants' " livestock ", such as aphids. When at rest, the ant-mimicking crab spider Amyciaea does not closely resemble Oecophylla , but while hunting it imitates the behavior of a dying ant to attract worker ants.
After a kill, some ant-mimicking spiders hold their victims between themselves and large groups of ants to avoid being attacked.
There is strong evidence that spiders' coloration is camouflage that helps them to evade their major predators, birds and parasitic wasps , both of which have good color vision.
Many spider species are colored so as to merge with their most common backgrounds, and some have disruptive coloration , stripes and blotches that break up their outlines.
In a few species, such as the Hawaiian happy-face spider, Theridion grallator , several coloration schemes are present in a ratio that appears to remain constant, and this may make it more difficult for predators to recognize the species.
Most spiders are insufficiently dangerous or unpleasant-tasting for warning coloration to offer much benefit. However, a few species with powerful venom, large jaws or irritant bristles have patches of warning colors, and some actively display these colors when threatened.
Many of the family Theraphosidae , which includes tarantulas and baboon spiders , have urticating hairs on their abdomens and use their legs to flick them at attackers.
These bristles are fine setae bristles with fragile bases and a row of barbs on the tip. The barbs cause intense irritation but there is no evidence that they carry any kind of venom.
A few spider species that build webs live together in large colonies and show social behavior, although not as complex as in social insects.
Anelosimus eximius in the family Theridiidae can form colonies of up to 50, individuals. For example, although Theridion nigroannulatum belongs to a genus with no other social species, T.
There is no consistent relationship between the classification of spiders and the types of web they build: species in the same genus may build very similar or significantly different webs.
Nor is there much correspondence between spiders' classification and the chemical composition of their silks. Convergent evolution in web construction, in other words use of similar techniques by remotely related species, is rampant.
Orb web designs and the spinning behaviors that produce them are the best understood. The basic radial-then-spiral sequence visible in orb webs and the sense of direction required to build them may have been inherited from the common ancestors of most spider groups.
It used to be thought that the sticky orb web was an evolutionary innovation resulting in the diversification of the Orbiculariae. Their greater success may be because sphecid wasps , which are often the dominant predators of spiders, much prefer to attack spiders that have flat webs.
About half the potential prey that hit orb webs escape. A web has to perform three functions: intercepting the prey intersection , absorbing its momentum without breaking stopping , and trapping the prey by entangling it or sticking to it retention.
No single design is best for all prey. For example: wider spacing of lines will increase the web's area and hence its ability to intercept prey, but reduce its stopping power and retention; closer spacing, larger sticky droplets and thicker lines would improve retention, but would make it easier for potential prey to see and avoid the web, at least during the day.
However, there are no consistent differences between orb webs built for use during the day and those built for use at night. In fact, there is no simple relationship between orb web design features and the prey they capture, as each orb-weaving species takes a wide range of prey.
The hubs of orb webs, where the spiders lurk, are usually above the center, as the spiders can move downwards faster than upwards. If there is an obvious direction in which the spider can retreat to avoid its own predators, the hub is usually offset towards that direction.
Horizontal orb webs are fairly common, despite being less effective at intercepting and retaining prey and more vulnerable to damage by rain and falling debris.
Various researchers have suggested that horizontal webs offer compensating advantages, such as reduced vulnerability to wind damage; reduced visibility to prey flying upwards, because of the backlighting from the sky; enabling oscillations to catch insects in slow horizontal flight.
However, there is no single explanation for the common use of horizontal orb webs. Spiders often attach highly visible silk bands, called decorations or stabilimenta, to their webs.
Field research suggests that webs with more decorative bands captured more prey per hour. There are several unusual variants of orb web, many of them convergently evolved, including: attachment of lines to the surface of water, possibly to trap insects in or on the surface; webs with twigs through their centers, possibly to hide the spiders from predators; "ladderlike" webs that appear most effective in catching moths.
However, the significance of many variations is unclear. In , Skylab 3 took two orb-web spiders into space to test their web-spinning capabilities in zero gravity.
At first, both produced rather sloppy webs, but they adapted quickly. Members of the family Theridiidae weave irregular, tangled, three-dimensional webs, popularly known as cobwebs.
There seems to be an evolutionary trend towards a reduction in the amount of sticky silk used, leading to its total absence in some species.
The construction of cobwebs is less stereotyped than that of orb-webs, and may take several days. The Linyphiidae generally make horizontal but uneven sheets, with tangles of stopping threads above.
Insects that hit the stopping threads fall onto the sheet or are shaken onto it by the spider, and are held by sticky threads on the sheet until the spider can attack from below.
Although the fossil record of spiders is considered poor,  almost species have been described from fossils.
In addition to preserving spiders' anatomy in very fine detail, pieces of amber show spiders mating, killing prey, producing silk and possibly caring for their young.
Hence Attercopus and the similar Permian arachnid Permarachne may not have been true spiders, and probably used silk for lining nests or producing egg cases rather than for building webs.
Several Carboniferous spiders were members of the Mesothelae , a primitive group now represented only by the Liphistiidae.
Some Triassic mygalomorphs appear to be members of the family Hexathelidae , whose modern members include the notorious Sydney funnel-web spider , and their spinnerets appear adapted for building funnel-shaped webs to catch jumping insects.
Araneomorphae account for the great majority of modern spiders, including those that weave the familiar orb-shaped webs.
The Jurassic and Cretaceous periods provide a large number of fossil spiders, including representatives of many modern families. The spiders Araneae are monophyletic i.
Arachnids lack some features of other chelicerates, including backward-pointing mouths and gnathobases "jaw bases" at the bases of their legs;  both of these features are part of the ancestral arthropod feeding system.
Pycnogonida sea spiders. Xiphosura horseshoe crabs. Araneae spiders. Pedipalpi whip scorpions , etc. The cladogram shows the relation among spider suborders and families: .
Spiders are divided into two suborders, Mesothelae and Opisthothelae , of which the latter contains two infraorders, Mygalomorphae and Araneomorphae.
Over 48, living species of spiders order Araneae have been identified and as of grouped into families and about 4, genera by arachnologists.